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Perspective Ecology
What makes a terrestrial ecosystem resilient?

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02 Mar 2018: Vol. 359, Issue 6379, pp. 988-989 DOI: 10.1126/science.aar5439
Kathy J. Willis
Royal Botanic Gardens, Kew, Richmond TW9 3AB, UK. Oxford Long-term Ecology Laboratory, Department of Zoology, University of Oxford, Oxford OX1 3PS, UK.
Elizabeth S. Jeffers
Oxford Long-term Ecology Laboratory, Department of Zoology, University of Oxford, Oxford OX1 3PS, UK.
Carolina Tovar
Royal Botanic Gardens, Kew, Richmond TW9 3AB, UK.

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With increasing incidence of extreme climatic events, disease outbreaks, and other environmental perturbations, conservation of terrestrial ecosystems that can retain their structure and function despite environmental shocks has moved rapidly up the international political agenda. International environmental policies and targets such as the Aichi Biodiversity Targets and the Sustainable Development Goals include conserving resilient ecosystems as a key priority.


This is an article distributed under the terms of the Science Journals Default License .


Vol 359, Issue 637902 March 2018

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What makes a terrestrial ecosystem resilient?

By Kathy J. Willis , Elizabeth S. Jeffers , Carolina Tovar

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Unitas surged back in 1964 and won his second MVP 15x28x39 cm (B x H T) Taschendieb Td0111ol Td0111ol Womens CrossBody Bag Brown 15x28x39 cm B x H T Taschendieb Womens Cross-Body Bag Brown modno14
while returning an improved Colts team to the playoffs with a 12-2 record. Baltimore would go 40-9-3 over this stretch with Unitas at the helm, and he would add a third MVP title to his collection in 1967. Unitas led the league in interceptions in 1966, but nobody was a credible threat to his title over this four-year stretch. Unitas’s second run produced a series of great teams, and one more title in 1970. 3

The Colts were the NFL’s representative in Super Bowl III after the 1968 season, but it was Unitas’s backup, Earl Morrall, at the helm most of the way. Unitas reentered the lineup when Morrall struggled against the Jets, but he wasn’t able to bring the Colts back. He was restored to the starting lineup in 1969, but a 36-year-old Unitas simply wasn’t the same player after an arm injury, and he was never a threat to hold this title again.

Reign : 1968-69

Namath is one of those players who inspire a lot of overrated quips by researchers who look back and see some pretty mediocre statistics coupled with injury issues and a career built on a famous quote. I used to be one of those very people, but I’ve been convinced otherwise by smart people like Chase Stuart, who did a great job of putting Namath’s career in context . He was first-team everything in 1968 even before the Super Bowl victory, and there was no NFL quarterback who matched his AFL reputation. He won the AFL AP Player of the Year award in 1969, a season when he was probably about equal to Raiders quarterback Daryle Lamonica. A year later, the two leagues would merge, but Namath missed most of the season with an injury, and Lamonica was overshadowed by another Bay Area passer …

Reign : 1970

Brodie was the best quarterback in the newly merged league by a pretty significant margin in 1970; he led the NFL in completions, passing yards, passing touchdowns, and had the lowest interception rate. He was first-team All-Pro and won the MVP award. It was a fluke season for the 35-year-old, and he would be San Francisco’s starter for only another year and a half after the season, but Brodie was brilliant in ’70.

Reign : 1971-73

The best young quarterback in the league had been Griese, who made the Pro Bowl in each of his first two seasons as a starter before becoming an MVP candidate in 1971. In leading Miami to the Super Bowl, Griese was the first-team All-Pro quarterback, finishing ahead of Roger Staubach in virtually every poll between the two. Griese would get hurt during Miami’s perfect season in 1972, but he would eventually take over during the AFC Championship Game for his scuffling replacement (Morrall again, perhaps not coincidentally) and lead Miami to consecutive wins for Don Shula’s first Super Bowl win as head coach. A healthy Griese would lead a dominant Dolphins team back to the promised land in 1973 before dropping off to above-average play for the next several seasons.


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